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Zoology and Entomology

Staff Member

Dr (CR) Charles Haddad
Senior Lecturer
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Short CV

PhD (Entomology) - University of the Free State, 2012



Publications (Short List)

Publications (2012-2014)



HADDAD, C.R. 2012. A revision of the spider genus Echinax Deeleman-Reinhold, 2001 (Araneae: Corinnidae) in the Afrotropical Region. Zootaxa 3450: 33–61.

HADDAD, C.R. 2012. A revision of the Afrotropical spider genus Cambalida Simon, 1909 (Araneae: Corinnidae). ZooKeys 234: 67–119.

HADDAD, C.R. & LOUW, S.vdM. 2012. A redescription of Merenius alberti Lessert, 1923 (Araneae: Corinnidae), with remarks on colour polymorphism and its relationship to ant models. African Invertebrates 53: 571–591.

ŠTÁHLAVSKÝ, F., KRÁL, J., HARVEY, M.S. & HADDAD, C.R. 2012. The first cytogenetic characterisation of atemnids: pseudoscorpions with the highest chromosome numbers (Arachnida: Pseudoscorpiones). Cytogenetic and Genome Research 137: 22–30.




DIPPENAAR-SCHOEMAN, A.S., VAN DEN BERG, A.M., HADDAD, C.R. & LYLE R. 2013. Current knowledge of spiders in South African agroecosystems (Arachnida, Araneae). Transactions of the Royal Society of South Africa 68: 57–74.

DIPPENAAR-SCHOEMAN, A.S., VAN DEN BERG, A.M., LYLE, R., HADDAD, C.R., FOORD, S.H & LOTZ, L.N. 2013. Die diversiteit van Suid-Afrikaanse spinnekoppe (Arachnida: Araneae): dokumentering van ‘n nasionale opname. Tydskrif van die Suid-Afrikaanse Akademie vir Wetenskap en Kuns 32(#375): 1–7.

FOURIE, R., HADDAD, C.R., DIPPENAAR-SCHOEMAN, A.S. & GROBLER, A. 2013. Ecology of the plant-dwelling spiders (Arachnida: Araneae) of the Erfenis Dam Nature Reserve, South Africa. Koedoe 55(#1113): 1–9.

HADDAD, C.R. 2013. A revision of the continental species of Copa Simon, 1885 (Araneae:Corinnidae) in the Afrotropical Region. ZooKeys 276: 1–37.

HADDAD, C.R. 2013. A revision of the ant-like sac spider genus Apochinomma Pavesi, 1881 (Araneae: Corinnidae) in the Afrotropical Region. Journal of Natural History 47: 2493–2529.

HADDAD, C.R. 2013. Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae. Zootaxa 3688: 1–79.

HADDAD, C.R., DIPPENAAR-SCHOEMAN, A.S., FOORD, S.H., LOTZ, L.N. & LYLE, R. 2013. The faunistic diversity of spiders (Arachnida: Araneae) of the Grassland Biome in South Africa. Transactions of the Royal Society of South Africa 68: 97–122.

HADDAD, C.R. & BOSMANS, R. 2013. Synonymy of the North African spider genus Castanilla Caporiacco, 1936 with Micaria Westring, 1851 (Araneae: Gnaphosidae). Zootaxa 3734: 397–399.

HADDAD, C.R. & WESOLOWSKA, W. 2013. Additions to the jumping spider fauna of South Africa (Araneae: Salticidae). Genus 24: 459–501.

KRÁL, J., KORÍNKOVÁ, T., KRKAVCOVÁ, L., MUSILOVÁ, J., ÁVILA HERRERA, I.M., FORMAN, M., VÍTKOVÁ, M., HADDAD, C.R., HEDIN, M., HENRIQUES, S. & PALACIOS VARGAS, J.G. 2013. Evolution of karyotype, sex chromosomes, and meiosis in mygalomorph spiders (Araneae: Mygalomorphae). Biological Journal of the Linnean Society 109: 377–408.

NEETHLING, J.A. & HADDAD, C.R. 2013. Arboreal spider assemblages associated with four tree species in the Grassland Biome of central South Africa (Arachnida: Araneae). Transactions of the Royal Society of South Africa 68: 123–131.

WESOLOWSKA, W. & HADDAD, C.R. 2013. New data on the jumping spiders of South Africa (Araneae: Salticidae). African Invertebrates 54: 177–240.




HADDAD, C.R. 2014. A revision of the endemic South African sac spider genus Lessertina Lawrence, 1942 (Araneae: Eutichuridae). Zootaxa 3873: 82–92.

RAMÍREZ, M.J., GRISMADO, C.J., LABARQUE, F.M., IZQUIERDO, M.A., LEDFORD, J.M., MILLER, J.A., HADDAD, C.R. & GRISWOLD, C.E. 2014. The morphology and relationships of the walking mud spiders of the genus Cryptothele (Araneae: Zodariidae). Zoologischer Anzeiger 253: 382–393.

WESOLOWSKA, W. & HADDAD, C.R. 2014. An overview of the jumping spiders of Lesotho (Araneae: Salticidae), with descriptions of six new species. African Invertebrates 55: 229–268.


Area(s) of Interest

African Arachnology. Various aspects of arachnid systematics and ecology are studied. The main focus is on the systematics and ecology of Afrotropical corinnid sac spiders, particularly ant-mimics in the subfamily Castianeirinae. This is one of the most poorly studied spider families in the region, and generic revisions presently underway indicate that the total diversity of this group is likely to increase at least five-fold. 

Other research areas include studies of spider biodiversity in habitats in the Free State Province, agroecosystems, and various conservancies. Biodiversity and ecological work focuses on the spider diversity of the Ndumo Game Reserve in Maputaland, KwaZulu-Natal, where more than 430 species of spiders have been recorded so far. Work on the role of spiders as predators of various insect pests in crops, and of termites in grassland, also receives attention. Collaborative research includes studies on the impacts of pesticides on spiders in crops, ecology of termitophagous and myrmecophagous spiders, and karyology of South African arachnids.

Courses Presented

ENT 226 – Introductory insect physiology (Sole lecturer). 
ENT 694 – Arachnology capita selecta (Sole lecturer).