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Zoology and Entomology

Staff Member

Dr (CR) Charles Haddad
Senior Lecturer
D 208
IB 49
051 4012568
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Short CV

PhD (Entomology) - University of the Free State, 2012



Publications (Short List)

Publications (2010-2012)


HADDAD, C.R. 2010. A new species of Poachelas from Maputaland, South Africa (Araneae: Corinnidae), with considerable range extension for Poachelas striatus. African Invertebrates 51: 313–319.

HADDAD, C.R. & BOSSELAERS, J. 2010. A revision of the genus Medmassa Simon, 1887 (Araneae: Corinnidae) in the Afrotropical Region. Zootaxa 2361: 1–12.

HADDAD, C.R., HONIBALL, A.S., DIPPENAAR-SCHOEMAN, A.S., SLOTOW, R. & VAN RENSBURG, B.J. 2010. Spiders (Arachnida: Araneae) as indicators of elephant-induced habitat changes in the Maputaland Centre of Endemism, South Africa. African Journal of Ecology 48: 446–460. 

HADDAD, C.R. & RUSSELL-SMITH, A. 2010. A comparison of spider diversity patterns in the Mkomazi Game Reserve, Tanzania and the Ndumo Game Reserve, South Africa (Arachnida: Araneae). African Journal of Ecology 48: 418–427.

LYLE, R. & HADDAD, C.R. 2010. A revision of the tracheline sac spider genus Cetonana Strand, 1929 in the Afrotropical Region, with descriptions of two new genera (Araneae: Corinnidae). African Invertebrates 51: 321–384.

MILLER, J.A., CARMICHAEL, A., RAMIREZ, M.J., HADDAD, C.R., REZÁC, M., JOHANNESEN, J., KRAL, J., WANG, X.P., SPAGNA, J.C. & GRISWOLD, C.E. 2010. Phylogeny of entelegyne spiders: affinities of the family Penestomidae (NEW RANK), generic phylogeny of Eresidae, and asymmetric rates of change in spinning organ evolution (Araneae, Araneoidea, Entelegynae). Molecular Phylogenetics and Evolution 55: 786–804.

MILLER, J.A., GRISWOLD, C.E. & HADDAD, C.R. 2010. Taxonomic revision of the spider family Penestomidae (Araneae, Entelegynae). Zootaxa 2534: 1–36.



BUTLER, V.P. & HADDAD, C.R. 2011. Spider assemblages associated with leaf litter of three tree species in central South Africa (Arachnida: Araneae). African Journal of Ecology 49: 301–310.

DIPPENAAR-SCHOEMAN, A.S., HAMER, M. & HADDAD, C.R. 2011. An annotated checklist of the spiders (Arachnida: Araneae) of the Mkambathi Nature Reserve, Eastern Cape, South Africa. Koedoe 53(#1058): 1–10. 

FOORD, S.H., DIPPENAAR-SCHOEMAN, A.S. & HADDAD, C.R. 2011. The faunistic diversity of spiders (Arachnida, Araneae) of the Savanna Biome in South Africa. Transactions of the Royal Society of South Africa 66: 170–201.

FOURIE, R., HADDAD, C.R. & JOCQUÉ, R. 2011. A revision of the purse-web spider genus Calommata Lucas, 1837 (Araneae: Atypidae) in the Afrotropical Region. ZooKeys 95: 1–28.

HADDAD, C.R., NEETHLING, J.A. & LYLE, R. 2011. Spinotrachelas montanus sp. n., the first Afromontane representative in the genus (Araneae: Corinnidae). African Invertebrates 52: 345–352.

HADDAD, C.R. & WESOLOWSKA, W. 2011. New species and new records of jumping spiders from central South Africa (Araneae: Salticidae). African Invertebrates 52: 51–134.

PEKAR, S. & HADDAD, C.R. 2011. Trophic strategy of ant-eating Mexcala elegans (Araneae: Salticidae): looking for evidence of evolution of prey-specialization. Journal of Arachnology 39: 133–138.



HADDAD, C.R. 2012. A revision of the spider genus Echinax Deeleman-Reinhold, 2001 (Araneae: Corinnidae) in the Afrotropical Region. Zootaxa 3450: 33–61.

HADDAD, C.R. 2012. A revision of the Afrotropical ant-like sac spider genus Cambalida Simon, 1909 (Araneae: Corinnidae). ZooKeys 234: 67–119.

HADDAD, C.R. & LOUW, S.vdM. 2012. A redescription of Merenius alberti Lessert, 1923 (Araneae: Corinnidae), with remarks on colour polymorphism and its relationship to ant models. African Invertebrates 53: 571–591.

ŠTÁHLAVSKÝ, F., KRÁL, J., HARVEY, M.S. & HADDAD, C.R. 2012. The first cytogenetic characterisation of atemnids: pseudoscorpions with the highest chromosome numbers (Arachnida: Pseudoscorpiones). Cytogenetic and Genome Research 137: 22–30.

Area(s) of Interest

African Arachnology. Various aspects of arachnid systematics and ecology are studied. The main focus is on the systematics and ecology of Afrotropical corinnid sac spiders, particularly ant-mimics in the subfamily Castianeirinae. This is one of the most poorly studied spider families in the region, and generic revisions presently underway indicate that the total diversity of this group is likely to increase at least five-fold. 

Other research areas include studies of spider biodiversity in habitats in the Free State Province, agroecosystems, and various conservancies. Biodiversity and ecological work focuses on the spider diversity of the Ndumo Game Reserve in Maputaland, KwaZulu-Natal, where more than 430 species of spiders have been recorded so far. Work on the role of spiders as predators of various insect pests in crops, and of termites in grassland, also receives attention. Collaborative research includes studies on the impacts of pesticides on spiders in crops, ecology of termitophagous and myrmecophagous spiders, and karyology of South African arachnids.

Courses Presented

ENT 226 – Introductory insect physiology (Sole lecturer). 
ENT 694 – Arachnology capita selecta (Sole lecturer).